1gene locus_value 2allele allele_value 3Locus_Tag locus_tag_value 4gene_syn synonym1 5gene-synonym synonym2 6protein-desc protein_descriptor 7protein protein_name1 8protein protein_name2 9function activity1 10activity activity2 11Ec_Number ec1 12EC-number ec2 13topology circular 14topology cirCular 15